Nivius

Nivius was the Makuta of Stelt.

Early History
He was created around 100,000 years ago with the other Makuta, and after that, everything that happened to most Makuta happened to him. A few thousand years ago, an inhabitant of his region, Sidorak, presented Nivius with his clan leader, Voporak for him to experiment on. Nivius converted Voporak into a hideous being sensitive to time shifts, and due to his unloyal nature, sent him to the Dark Hunters to learn discipline, but abandoned his experiment in fury after discovering Voporak had become loyal to the Dark Hunters and not the Brotherhood.

Post-Great Cataclysm
During the Great Cataclysm, part of the Destral fortress fell onto Nivius, crushing his armor. Mutran and Chirox managed to collect up his antidermis and place it inside an Exo-Toa body before his antidermis dissipated. He remained inside this armor suit until his own armor was repaired.

Karda Nui
He joined the Nexensis when it formed, and headed to Karda Nui with the other members on Teridax's orders. He was mutated into a vaguely reptilian creature by the swamp water, and lost several abilities. He later escaped Karda Nui with the other Nexensis, just before the Energy Storm began. They sought refuge in The Cavern of Sorrow, and swayed Atrox into joining their cause. The Nexensis now had a new objective: defeat Teridax in any way possible, no matter who had to die as a result. Nivius went with Atra, Rutilus and Viridis to destroy Karda Nui, while Atrox, Vitiosus and Crocinus headed to the Core Processor to keep Teridax distracted.